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I recall Markus say explicitly in a thread on 12 or 13 that rust was only in the offseason. I think rehab assignments are there (1) because they're an MLB reality and (2) they buy you time to figure out your roster changes.
Though, now that I think about it more, I think Markus said rust only happens in the offseason and with long-term (2 weeks plus, I think?) injuries. I do not think it happens if you sit a sub on the bench for 150 games then play him.
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